Organizational boundaries are critical for defining what is inside and outside an organization, enabling its recognition within the external environment. This research investigates the unique dynamics of free and open-source software (FOSS) development, where evolving value systems, external influences, and new collaboration models reshape already porous and ambiguous boundaries. Through an in-depth single case study, we explore how organizational practices adapt to boundary fluctuations. Our findings reveal that interacting, communicating, and aligning inter- and intra-organizational practices are closely tied to boundary dynamics. Values, technology, and money emerge as key drivers, shaping how practices are enacted and adapted and generating tensions within these practices. During boundary shifts, practices may remain stable, evolve, or diminish, depending on the dominance of specific drivers. Building on these insights, we explore the interplay between these elements and propose a framework that illustrates the interrelationship between boundary dynamics, practices, and drivers.